Lyra

Lyra begins as an alchemist under the command of Lieutenant Yoki in Youswell with aspirations of becoming a State Alchemist. Lyra'sl alchemic attacks condense the moisture in the air and force it at high velocity toward her adversary. Her necklace bears a small transmutation circle. After being defeated by Edward in a battle, she seems to learn from her mistake in following Yoki and becomes the apprentice of Dante three years later in an effort to improve her already strong abilities and become an even greater alchemist. Dante, whose body was aging, performed a body switch with Lyra using the Philosopher's Stone and then proceeded to kill the original Lyra so that her secret would remain hidden.

Edward later finds the remains of Dante's old body and assumes the homunculus Greed was responsible.